2023年6月24日发(作者：)

J.T. Eastman

Jan. 13, 2010

Table 1. Notothenioid classification and list of species. Arrangement is phylogenetic for families,

based on Near et al. (2004) and Near and Cheng (2008), and alphabetical for genera within families

and for species within genera. Superfamilies from Voskoboinikova (2004). Fishes of the Southern

Ocean (Gon and Heemstra, 1990) is starting point for species list with subsequent additions noted.

Bovichtids from Hardy (1988), Balushkin (1992) and Bravo et al. (1999); pseudaphritids and

eleginopids from Balushkin (1992, 2000); nototheniids from DeWitt et al. (1990), but following

Schneppenheim et al. (1994) for the Lepidonotothen squamifrons group, and with non-Antarctic

species from DeWitt (1966, 1970); harpagiferids from Hureau (1990); artedidraconids from Eakin

(1990) with revision of Pogonophryne following Balushkin and Eakin (1998); bathydraconids from

Gon (1990) and channichthyids from Iwami and Kock (1990). Asterisk (*) indicates non-Antarctic

species. Dagger (†) precedes names of species described or resurrected from synonymy after

publication of Fishes of the Southern Ocean in 1990. Species placed in synonymy since 1990 are

underlined in the list but not counted. Other changes since Eastman and Eakin (2000) indicated by

footnotes. Catalog of Fishes (Eschmeyer, 1998; Eschmeyer and Fricke, 2009, online version) used

for authorities and dates in cases of uncertainty. Total count = 131 species (104 Antarctic and 27 non-Antarctic) and 44 genera. Up to date as of January 13, 2010.

Superfamilies, families and species

Bovichtoidea

Bovichtidae (9)

Bovichtus

angustifrons Regan, 1913*

argentinus MacDonagh, 1931*a

chilensis Regan, 1913*

diacanthus (Carmichael, 1819)*

elongatus Hureau and Tomo, 1977a

oculus Hardy, 1988*

psychrolutes Günther, 1860*

variegatus Richardson, 1846*

veneris Sauvage, 1879*

Cottoperca trigloides (Forster, 1801)*b

†Halaphritis platycephala Last, Balushkin and Hutchins, 2002*c

Notothenioidea

Pseudaphritidae (1)

Pseudaphritis urvillii (Valenciennes in Cuvier and Valenciennes, 1832)*

Eleginopidae (1)

Eleginops maclovinus (Cuvier in Cuvier and Valenciennes, 1830)*

Nototheniidae (50)

Aethotaxis mitopteryx DeWitt, 1962

Cryothenia

†amphitreta Cziko and Cheng, 2006d

peninsulae Daniels, 1981

Dissostichus

eleginoides Smitt, 1898

mawsoni Norman, 1937

Gobionotothen

acuta (Günther, 1880)

†barsukovi Balushkin, 1991

gibberifrons (Lönnberg, 1905)

marionensis (Günther, 1880)

Gvozdarus svetovidovi Balushkin, 1989

Lepidonotothen

kempi (Norman, 1937)

larseni (Lönnberg, 1905)

macrophthalma (Norman, 1937)*

mizops (Günther, 1880)

nudifrons (Lönnberg, 1905)

squamifrons (Günther, 1880)

Notothenia

angustata Hutton, 1875*

2

Trematomus

bernacchii Boulenger, 1902

eulepidotus Regan, 1914

hansoni Boulenger, 1902

lepidorhinus (Pappenheim, 1911)

loennbergii Regan, 1913

newnesi Boulenger, 1902

nicolai (Boulenger, 1902)

pennellii Regan, 1914

scotti (Boulenger, 1907)

coriiceps Richardson, 1844

cyanobrancha Richardson, 1844

microlepidota Hutton, 1875*

rossii Richardson, 1844

3

Pagothenia

borchgrevinki (Boulenger, 1902)

brachysoma (Pappenheim, 1912)

Paranotothenia

†dewitti Balushkin, 1990

magellanica (Forster in Bloch and Schneider, 1801)

Patagonotothen

brevicauda (Lönnberg, 1905)*

canina (Smitt, 1897)*

cornucola (Richardson, 1844)*

elegans (Günther, 1880)*

guntheri (Norman, 1937)

jordani (Thompson, 1916)*

†kreffti Balushkin and Stehmann, 1993*

longipes (Steindachner, 1876)*

ramsayi (Regan, 1913)*

sima (Richardson, 1844)*

squamiceps (Peters, 1876)*

tessellata (Richardson, 1845)*

†trigramma (Regan, 1913)*e

†thompsoni Balushkin, 1993*

wiltoni (Regan, 1913) *

Pleuragramma antarcticum Boulenger, 1902 tokarevi Andriashev, 1978

vicarius Lönnberg, 1905

Harpagiferidae (11)

Harpagifer

†andriashevi Prirodina, 2000f

antarcticus Nybelin, 1947

bispinis (Schneider in Bloch and Schneider, 1801)

†crozetensis Prirodina, 2004g

georgianus Nybelin, 1947

kerguelensis Nybelin, 1947

†macquariensis Prirodina, 2000f

†nybelini Prirodina, 2002h

†permitini Neyelov and Prirodina, 2006i

palliolatus Richardson, 1845

spinosus Hureau, Louis, Tomo and Ozouf, 1980

Artedidraconidae (27)

Artedidraco

†glareobarbatus Eastman and Eakin, 1999

loennbergi Roule, 1913

mirus Lönnberg, 1905

orianae Regan, 1914

shackletoni Waite, 1911

skottsbergi Lönnberg, 1905

Dolloidraco longedorsalis Roule, 1913

Histiodraco velifer (Regan, 1914)

Pogonophryne

albipinna Eakin, 1981

barsukovi Andriashev, 1967

†bellingshausenensis Eakin, Eastman and Matallanas, 2008j

†cerebropogon Eakin and Eastman, 1998

curtilemma Balushkin, 1988

dewitti Eakin, 1988

dolichobranchiata Andriashev, 1967

†eakini Balushkin, 1999

4 †fusca Balushkin and Eakin, 1998

immaculata Eakin, 1981

lanceobarbata Eakin, 1987

macropogon Eakin, 1981

marmorata Norman, 1938

mentella Andriashev, 1967

†orangiensis Eakin and Balushkin, 1998

orcadensis Tomo, 1981

permitini Andriashev, 1967

phyllopogon Andriashev, 1967

platypogon Eakin, 1988

scotti Regan, 1914

†stewarti Eakin, Eastman and Near, 2009k

†squamibarbata Eakin and Balushkin, 2000

velifera Eakin, 1981

ventrimaculata Eakin, 1987

Bathydraconidae (16)

†Acanthodraco dewitti Skóra, 1995

Akarotaxis nudiceps (Waite, 1916)

Bathydraco

antarcticus Günther, 1878

joannae DeWitt, 1985

macrolepis Boulenger, 1907

marri Norman, 1938

scotiae Dollo, 1906

Cygnodraco mawsoni Waite, 1916

Gerlachea australis Dollo, 1900

Gymnodraco acuticeps Boulenger, 1902

Parachaenichthys

charcoti (Vaillant, 1906)

georgianus (Fischer, 1885)

Prionodraco evansii Regan, 1914

5 Psilodraco breviceps Norman, 1937

Racovitzia glacialis Dollo, 1900

Vomeridens infuscipinnis (DeWitt, 1964)

Channichthyidae (16)

Chaenocephalus aceratus (Lönnberg, 1906)

Chaenodraco wilsoni Regan, 1914

Champsocephalus

esox (Günther, 1861)

gunnari Lönnberg, 1905

Channichthys rhinoceratus Richardson, 1844

Chionobathyscus dewitti Andriashev and Neelov, 1978

Chionodraco

hamatus (Lönnberg, 1905)

myersi DeWitt and Tyler, 1960

rastrospinosus DeWitt and Hureau, 1979

Cryodraco

antarcticus Dollo, 1900

†atkinsoni Regan, 1914l

Dacodraco hunteri Waite, 1916

Neopagetopsis ionah Nybelin, 1947

Pagetopsis

macropterus (Boulenger, 1907)

maculatus Barsukov and Permitin, 1958

Pseudochaenichthys georgianus Norman, 1937

6 a7

Bravo et al. (1999) placed these two species in the synonymy of Bovichtus chilensis. Eastman and

Eakin (2000) were unaware of this paper when compiling their 2000 list.

bThis is a change from Eastman and Eakin (2000). We now follow Balushkin (1992, 2000) and

Eschmeyer and Fricke (2009) in considering Cottoperca gobio (Günther, 1861) as a junior synonym

of C. trigloides (Forster in Bloch and Schneider, 1801).

cLast et al. (2002).

dCziko and Cheng (2006).

eNot included in previous lists; the status of this species is uncertain but it is treated here as valid; it is

widely recognized by South American ichthyologists, e.g., García and Menni (1996) and Ojeda et al.

(2000) and has recently been collected off West Falkland Island (Paul Brickle, Fisheries Department,

Falkland Islands Government, personal communication, 2005).

fPrirodina (2000).

gPrirodina (2004).

hPrirodina (2002).

iNeyelov and Prirodina (2006).

jEakin et al. (2008).

kEakin et al. (2009).

lLa Mesa et al. (2002).

8

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